Ptomaphaginus

Ptomaphaginus
Nomenklatur
Ptomaphaginus Portevin, 1914
Populærnavn
mycelbiller^[1]
Klassifikasjon
Rike		Dyr
Rekke		Leddyr
Klasse		Insekter
Orden		Biller
Underorden		Polyphaga
Familie		Mycelbiller
Økologi
Antall arter:		ca. 100
Habitat:		på land, ofte i huler
Utbredelse:		Asia
Inndelt i

Ptomaphaginus er en slekt av mycelbiller som særlig finnes knyttet til sopp og åtsler.

Utseende

Små (2–4 millimeter), halvmatte, mørkbrunlige mycelbiller, kroppsformen nærmest dråpeformet med lange bein. Antennene er litt kølleformede og 11-leddete. De fem ytterste leddene danner en avlang klubbe, men det åttende leddet er mye mindre enn de andre og er vanligvis forkortet til en liten skive. Hodet er oftest kraftig innskåret bak fasettøynene, med en smal hals, men dette kan være vanskelig å se da billene gjerne trekker hodet inn under brystskjoldet. Hannen har utvidete føtter på forbeina.

Levevis

De fleste av artene lever i huler eller nede i jorda, og er gjerne blinde.

Utbredelse

De aller fleste av artene er utbredt i Sørøst-Asia.

Systematisk inndeling

Treliste

Ordenen biller, Coleoptera
- Underordenen Polyphaga
  - Overfamilien åtsel- og rovbiller, Staphylinoidea
    - Familien mycelbiller, Leiodidae Fleming, 1821
      - Underfamilien kadaverbiller, Cholevinae
        Stammen Ptomaphagini
        Slekten Ptomaphaginus Portevin, 1914

Liste over artene

Ptomaphaginus agostii Schilthuizen & Perreau, 2008^[2]
Ptomaphaginus alabensis Schilthuizen & Perreau, 2018^[3]
Ptomaphaginus anas Schilthuizen & Perreau, 2008
Ptomaphaginus angusticornis (Portevin, 1921)
Ptomaphaginus apiculatus Szymczakowski, 1975
Ptomaphaginus assimilis Szymczakowski, 1972
Ptomaphaginus baliensis Perreau, 1995
Ptomaphaginus bengalicola Perreau, 1991
Ptomaphaginus bryanti Jeannel, 1936
Ptomaphaginus bryantioides Schilthuizen & Perreau, 2008
Ptomaphaginus bucculentus Szymczakowski, 1974
Ptomaphaginus burckhardti Schilthuizen & Perreau, 2008
Ptomaphaginus caroli Schilthuizen & Perreau, 2008
Ptomaphaginus chapmani Peck, 1981 ^[4]
Ptomaphaginus cherrapunjeensis Perreau, 1991
Ptomaphaginus cilipes (Portevin, 1907)
Ptomaphaginus clibanarius Szymczakowski, 1970
Ptomaphaginus coronatus Szymczakowski, 1974
Ptomaphaginus dao Perkovsky, 2001
Ptomaphaginus depequkri Perreau, 1986
Ptomaphaginus flavicornis (Motschulsky, 1863)
Ptomaphaginus fornicatus Szymczakowski, 1972
Ptomaphaginus franki Perreau, 1993
Ptomaphaginus geigenmuellerae Perreau, 1993
Ptomaphaginus giachinoi Schilthuizen & Perreau, 2008
Ptomaphaginus gibberosus Szymczakowski, 1972
Ptomaphaginus giganteus Szymczakowski, 1980
Ptomaphaginus gonbe Nishikawa & Hayashi, 2016^[5]
Ptomaphaginus gracilis Schweiger, 1956
Ptomaphaginus grandis Schilthuizen & Perreau, 2018
Ptomaphaginus guangxiensis Wang & Zhou, 2015^[6]
Ptomaphaginus gutianshanicus Wang & Zhou, 2015
Ptomaphaginus hamatus Szymczakowski, 1975
Ptomaphaginus hanskii Schilthuizen & Perreau, 2018
Ptomaphaginus heterotrichus Szymczakowski, 1975
Ptomaphaginus honestus Szymczakowski, 1964
Ptomaphaginus isabellarosselini Schilthuizen, Njunjić & Perreau, 2018
Ptomaphaginus ishizuchiensis Perreau, 1996
Ptomaphaginus jacobsoni Szymczakowski, 1964
Ptomaphaginus kinabaluensis Schilthuizen & Perreau, 2008
Ptomaphaginus kinabatanganensis Njunjić, Schilthuizen & Perreau, 2018
Ptomaphaginus kosiensis Perreau, 1988
Ptomaphaginus kurbatovi Schilthuizen & Perreau, 2008
Ptomaphaginus lacertosus Szymczakowski, 1970
Ptomaphaginus laticornis Jeannel, 1936
Ptomaphaginus latimanus Schilthuizen & Perreau, 2008
Ptomaphaginus latipes (Pic, 1929)
Ptomaphaginus layangensis Schilthuizen & Perreau, 2018
Ptomaphaginus leucodon Szymczakowski, 1975
Ptomaphaginus lipsae Perreau & Lemaire, 2018
Ptomaphaginus loeblianus Schilthuizen & Perreau, 2008
Ptomaphaginus longitarsis Portevin, 1914
Ptomaphaginus louis Schilthuizen & Perreau, 2018
Ptomaphaginus luoi Wang & Zhou, 2015
Ptomaphaginus major Szymczakowski, 1972
Ptomaphaginus marshalli Schilthuizen & Perreau, 2018
Ptomaphaginus megalayanus Perreau, 1991
Ptomaphaginus microphallus Schilthuizen & Perreau, 2018
Ptomaphaginus minimus Perreau, 1998
Ptomaphaginus mirabilis Szymczakowski, 1972
Ptomaphaginus miyataorum Nishikawa, 2017
Ptomaphaginus muluensis Schilthuizen & Perreau, 2018
Ptomaphaginus murphyi Szymczakowski, 1970
Ptomaphaginus nanus Schilthuizen & Perreau, 2018
Ptomaphaginus newtoni Wang & Zhou, 2015
Ptomaphaginus nipponensis Perreau, 1996
Ptomaphaginus nitens Jeannel, 1936
Ptomaphaginus obtusus Szymczakowski, 1959
Ptomaphaginus okinawaensis Hoshina & Sugaya, 2003
Ptomaphaginus oribates Szymczakowski, 1965
Ptomaphaginus pallidicornis (Portevin, 1907)
Ptomaphaginus palpalis Szymczakowski, 1974
Ptomaphaginus palpaloides Perreau, 1988
Ptomaphaginus pecki Perkovsky, 1995
Ptomaphaginus perreaui Wang & Zhou, 2015
Ptomaphaginus pilipennis Perreau, 1991
Ptomaphaginus pilipennoides Perreau, 1991
Ptomaphaginus pingtungensis Perreau, 1996
Ptomaphaginus piraster Szymczakowski, 1975
Ptomaphaginus portevini Szymczakowski, 1964
Ptomaphaginus quadricalcarus Wang & Zhou, 2015
Ptomaphaginus riedeli Perreau, 1995
Ptomaphaginus rubidus (Champion, 1927)
Ptomaphaginus rufus Jeannel, 1936
Ptomaphaginus rugosus Perreau, 1991
Ptomaphaginus ruzickai Wang & Zhou, 2015
Ptomaphaginus sabahensis Schilthuizen & Perreau, 2008
Ptomaphaginus sarawacensis Schilthuizen & Perreau, 2018
Ptomaphaginus sauteri (Portevin, 1914)
Ptomaphaginus scaber Szymczakowski, 1964
Ptomaphaginus scaphaner Szymczakowski, 1972
Ptomaphaginus schawalleri Perreau, 1992
Ptomaphaginus shennongensis Wang & Zhou, 2015
Ptomaphaginus shibatai Hayashi, 1969
Ptomaphaginus similipes Schilthuizen & Perreau, 2008
Ptomaphaginus similis Schweiger, 1956
Ptomaphaginus sinuatus Schilthuizen, 1984
Ptomaphaginus smetanai Perreau, 1988
Ptomaphaginus takaosanus Nakane, 1982
Ptomaphaginus takashii Nishikawa, 2017
Ptomaphaginus tarsalis Szymczakowski, 1964
Ptomaphaginus testaceus Schilthuizen & Perreau, 2018
Ptomaphaginus thieleni Perreau, 1993
Ptomaphaginus tomellerii Zoia, 1997
Ptomaphaginus trautneri Perreau, 1993
Ptomaphaginus troglodytes Perreau & Růžička, 2018^[7]
Ptomaphaginus truncatus Perreau, 1988
Ptomaphaginus turensis Perreau, 1991
Ptomaphaginus wenboi Wang & Zhou, 2015
Ptomaphaginus wuzhishanicus Wang & Zhou, 2015
Ptomaphaginus yaeyamaensis Hoshina & Sugaya, 2003
Ptomaphaginus yui Wang & Zhou, 2015

Referanser

^ «Artsdatabankens artsopplysninger». Artsdatabanken. 12. november 2020. Besøkt 12. november 2020.
^ Schilthuizen, Menno & Michel Perreau (2008). New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae). Zoologische Mededelingen 82 (1): 189-210.
^ Schilthuizen, Menno, Michel Perreau & Iva Njunjić (2018) A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae). ZooKeys 777: 57-108.
^ Peck, Stewart B (1981). «A new cave‐inhabiting Ptomaphaginus beetle from Sarawak». Systematic Entomology. 6 (2): 221–224. doi:10.1111/j.1365-3113.1981.tb00436.x.
^ Nishikawa, M. & Y. Hayashi (2016) A new Ptomaphaginus (Coleoptera, Leiodidae, Cholevinae) from Nagano Prefecture, Central Japan. Elytra, (N.S.) 6 (2): 347-351.
^ Wang, Cheng-Bin & Hong-Zhang Zhou (2015) Taxonomy of the genus Ptomaphaginus Portevin (Coleoptera: Leiodidae: Cholevinae) from China, with description of eleven new species. Zootaxa 3941 (3): 301-338.
^ Perreau, Michel; Růžička, Jan (2018). «Ptomaphaginus troglodytes sp. n., the first anophthalmic species of Ptomaphaginina from China». ZooKeys (749): 135–147. PMC 5904491 . PMID 29674925. doi:10.3897/zookeys.749.24964.